JACKSON, Wyo. — Grizzly 399, beloved bear with an international fan club, died on Tuesday night in a vehicle collision south of Jackson.

Buckrail photographer Nick Sulzer captured beautiful photos and videos of 399 over the last few years as she and her cubs traveled in and around Grand Teton National Park.

Video: Nick Sulzer // Buckrail

399’s bumper crop of four cubs made for fairytale-like photos.

Photo: Nick Sulzer // Buckrail

Considered the great matriarch of the Tetons, Grizzly 399 leaves behind a storied legacy.

Photo: Nick Sulzer // Buckrail
Video: Nick Sulzer // Buckrail

“Grizzly bear 399 has been perhaps the most prominent ambassador for the species,” Grand Teton National Park Superintendent Chip Jenkins said in the announcement of her death. “She has inspired countless visitors into conservation stewardship around the world and will be missed.”

Photo: Nick Sulzer // Buckrail
Photo: Nick Sulzer // Buckrail

Got a great photo of 399? Send it to tips@buckrail.com for the chance to be featured in an upcoming Snapped.

Video: Nick Sulzer // Buckrail

Marianne is the Editor of Buckrail. She handles breaking news and reports on a little bit of everything. She's interested in the diversity of our community, arts/entertainment and crazy weather.